Physalaemus guayaco Ron, Coloma & Cannatella 2005

Ron, Coloma & Cannatella 2005. A new, cryptic species of Physalaemus (Anura: Leptodactylidae) from western Ecuador with comments on the call structure of the P. pustulosus species group. Herpetologica, 61(2), June 2005: 178-198