Chandler 1992. Short-winged mould beetles of the tribe Arhytodini of Panama, with descriptions of new taxa (Coleoptera: Pselaphidae: Pselaphinae). Quintero, D.; Aiello, A.[Eds]. Insects of Panama and Mesoamerica: selected studies., Oxford University Press, Oxford, New York & Tokyo, 1992: i-xxii, 1-692. Chapter pagination: 339-344, 651, 660