The Taygetis ypthima species group (Lepidoptera, Nymphalidae, Satyrinae): taxonomy, variation and description of a new species

doi:10.3897/zookeys.356.6481

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Title	The Taygetis ypthima species group (Lepidoptera, Nymphalidae, Satyrinae): taxonomy, variation and description of a new species
Author(s)	Ricardo Siewert Thamara Zacca Fernando Dias André Freitas Olaf Mielke Mirna Casagrande
Journal	ZooKeys
ISSN	1313-2989
Volume	356
Pages	11--30
DOI	10.3897/zookeys.356.6481
Year	2013
Publisher	Pensoft Publishers
Citation

...

Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) de Matas Paludosas e Matas de Restinga da Planície Costeira da região Sul do Brasil.
by JM Bellaver;CA Iserhard;JP Santos;AK Silva;M Torres;RR Siewert;A Moser;HP Romanowski
Biota Neotropica 12 pages 181--190 (2012)
- 10.1590/S1676-06032012000400019
Arquivos de Entomologia, série A
by CM Biezanko
Butterflies (Lepidoptera: Papilionoidea e Hesperioidea) from two forest fragments in northern Rio Grande do Sul, Brazil.
by D Bonfantti;Mare RA Di;R Giovenardi
Check List 5 pages 819--829 (2009)
Chromosomal evolution of South American frugivorous butterflies in the Satyroid clade (Nymphalidae: Charaxinae, Morphinae and Satyrinae).
by Jr KS Brown;AVL Freitas;BV Schoultz;AO Saura;A Saura
Biological Journal of the Linnean Society 92 pages 467--481 (2007)
- 10.1111/j.1095-8312.2007.00872.x
by B D’Abrera
Borboletas (Papilionoidea e Hesperioidea) de Guarapuava e arredores, Paraná, Brasil: um inventário com base em 63 anos de registros.
by DR Dolibaina;OHH Mielke;MM Casagrande
Biota Neotropica 11 pages 343--354 (2011)
- 10.1590/s1676-06032011000100031
On the frequency of butterflies in eastern Brazil, with a list of butterfly fauna of Poços de Caldas, Minas Gerais.
by H Ebert
Journal of the Lepidopterists' Society 23 pages 1--48 (1969)
- 144150
Beiträge zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae.
by W Forster
Veröffentlichungen der zoologischen Staatssammlung München 8 pages 51--188 (1964)
- 113851
A new species of Yphthimoides (Nymphalidae, Satyrinae) from Southeastern Brazil.
by AVL Freitas
Journal of the Lepidopterists' Society 58 pages 7--12 (2004)
- 113839
A new species of Moneuptychia Forster (Lepidoptera: Satyrinae, Euptychiina) from the highlands of southeastern Brazil.
by AVL Freitas
Neotropical Entomology 36 pages 919--925 (2007)
- 10.1590/s1519-566x2007000600014
Description of genus Guaianaza for “Euptychia” pronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages.
by AVL Freitas;C Peña
Zootaxa 1163 pages 49--59 (2006)
A new species of Moneuptychia Forster (Lepidoptera: Satyrinae: Euptychiina) from central Brazil.
by AVL Freitas;EO Emery;OHH Mielke
Neotropical Entomology 39 pages 83--90 (2010)
- 10.1590/s1519-566x2010000100011
A new genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from southern Brazil.
by AVL Freitas;OHH Mielke;A Moser;KL Silva-Brandão;CA Iserhard
Neotropical Entomology 40 pages 231--237 (2011)
- 10.1590/S1519-566X2011000200012
Zoologia, Curitiba.
by AVL Freitas;EP Barbosa;JP Santos;OHH Mielke
Diversidade de Lepidoptera (Papilionoidea e Hesperioidea) em dois fragmentos de floresta no município de Frederico Westphalen, Rio Grande do Sul, Brasil.
by R Giovenardi;RA Di Mare;J Sponchiado;S Roani;FAF Jacomassa;AB Jung;MA Porn
Revista Brasileira de Entomologia 52 pages 599--605 (2008)
- 10.1590/S0085-56262008000400010
Lista de espécies de borboletas (Lepidoptera, Papilionoidea e Hesperioidea) da região do vale do rio Maquiné, Rio Grande do Sul, Brasil.
by CA Iserhard;HP Romanowski
Revista Brasileira de Zoologia 21 pages 649--662 (2004)
- 10.1590/s0101-81752004000300027
Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) ocorrentes em diferentes ambientes na Floresta Ombrófila Mista e nos Campos de Cima da Serra do Rio Grande do Sul, Brasil.
by CA Iserhard;MT Quadros;HP Romanowski;MS Mendonça
Biota Neotropica 10 pages 309--320 (2010)
- 10.1590/s1676-06032010000100026
by JA Kochalka;D Torres;B Garcete;C Aguilar
Papilionoidea (Lepidoptera) de Pelotas e seus arredores, Rio Grande do Sul, Brasil.
by CP Krüger;EJE Silva
Entomologia y Vectores 10 pages 31--45 (2003)
by G Lamas
From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera: Nymphalidae): history, progress and prospects.
by MA Marín;C Peña;AVL Freitas;N Wahlberg;SI Uribe
Neotropical Entomology 40 pages 1--13 (2011)
- 10.1590/s1519-566x2011000100001
Systematics and evolutionary history of butterflies in the ‘‘Taygetis clade’'(Nymphalidae: Satyrinae: Euptychiina): Towards a better understanding of Neotropical biogeography.
by PF Matos-Maraví;C Peña;KR Willmott;AVL Freitas;N Wahlberg
Molecular Phylogenetics and Evolution 66 pages 54--68 (2013)
- 10.1016/j.ympev.2012.09.005
by DL Murray
Immature stages and biology of Taygetis Hübner (Lepidoptera: Nymphalidae).
by DL Murray
Proceedings of the Entomological Society of Washington 103 pages 932--945 (2001b)
A catalogue of the type specimens of Nymphalidae deposited in the collection of the Staatliches Museum für Tierkunde Dresden (Insecta: Lepidoptera: Rhopalocera).
by YP Nekrutenko
Entomologische Abhandlungen Museum für Tierkunde Dresden 59 pages 325--403 (2001)
Bulletin de la Sociéte Entomologique de Mulhouse
by EV Niculescu
Sobre a morfologia do penis em Lepidoptera.
by J Oiticica-Filho
Boletim do Museu Nacional, Rio de Janeiro, n. s., Zoologia 50 pages 1--79 (1946)
Nymphalidae, Papilionidae e Pieridae (Lepidoptera: Papilionoidea) da Serra do Sudoeste do Rio Grande do Sul, Brasil.
by ALG Paz;HP Romanowski;ABB Morais
Biota Neotropica 8 pages 141--149 (2008)
- 10.1590/s1676-06032008000100017
Occurrence of fruit-feeding butterflies (Lepidoptera: Nymphalidae) in a fragment of Araucaria Moist Forest in Rio Grande do Sul State, Brazil.
by VS Pedrotti;MP Barros;HP Romanowski;CA Iserhard
Biota Neotropica 11 pages 385--390 (2011)
- 10.1590/S1676-06032011000100036
Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae).
by C Peña;G Lamas
Revista Peruana de Biologia 12 pages 5--48 (2005)
Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae).
by C Peña;S Nylin;AVL Freitas;N Wahlberg
Zoologica Scripta 39 pages 243--258 (2010)
- 10.1111/j.1463-6409.2010.00421.x
Ninfalídeos (Lepidoptera, Nymphalidae) ocorrentes no norte da Planície Costeira do Rio Grande do Sul, Brasil.
by FC Quadros;AL Dorneles;E Corseuil
Biociências 12 pages 147--164 (2004)
The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape.
by DB Ribeiro;R Batista;PI Prado;Jr. KS Brown;AVL Freitas
Biodiversity and Conservation 21 pages 811--827 (2012)
- 10.1007/s10531-011-0222-x
Fruit-feeding butterflies guide of subtropical Atlantic Forest and Araucaria Moist Forest in State of Rio Grande do Sul, Brazil.
by JP Santos;CA Iserhard;MO Teixeira;HP Romanowski
Biota Neotropica 11 pages 253--274 (2011)
- 10.1590/s1676-06032011000300022
Borboletas frugívoras (Lepidoptera: Nymphalidae) de uma área urbana (Área de Proteção Especial Manancial Cercadinho) em Belo Horizonte, Minas Gerais, Brasil.
by ARM Silva;CO Castro;PO Mafia;MOC Mendonça;TCC Alves;MV Beirão
Biota Neotropica 12 pages 292--297 (2012)
- 10.1590/S1676-06032012000300028
Preliminary analysis of the diurnal Lepidoptera fauna of the Três Picos State Park, Rio de Janeiro, Brazil, with a note on Parides ascanius (Cramer, 1775).
by A Soares;JMS Bizarro;CB Bastos;N Tangerini;NA Silva;AS Silva;GB Silva
Tropical Lepidoptera Research 21 pages 66--79 (2011)
Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (São Paulo).
by M Uehara-Prado;AVL Freitas;RB Francini;Jr KS Brown
Biota Neotropica 4 pages 1--5 (2004)
- 10.1590/S1676-06032004000100007
Illustrated Lists of American Butterflies.
by AD Warren;KJ Davis;EM Stangeland;JP Pelham;NV Grishin
Stegosatyrus, a new genus of Euptychiina from the grasslands of neotropical realm (Lepidoptera: Nymphalidae: Satyrinae).
by T Zacca;OHH Mielke;TW Pyrcz;MM Casagrande;AVL Freitas;P Boyer
Zootaxa 3682 pages 331--350 (2013)
- 10.11646/zootaxa.3682.2.7

Only known citations between publications in BioNames are listed here.

[1] Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) de Matas Paludosas e Matas de Restinga da Planície Costeira da região Sul do Brasil.
by JM Bellaver;CA Iserhard;JP Santos;AK Silva;M Torres;RR Siewert;A Moser;HP Romanowski
Biota Neotropica 12 pages 181--190 (2012)
10.1590/S1676-06032012000400019

[2] 10.1590/S1676-06032012000400019

[3] Arquivos de Entomologia, série A
by CM Biezanko

[4] Butterflies (Lepidoptera: Papilionoidea e Hesperioidea) from two forest fragments in northern Rio Grande do Sul, Brazil.
by D Bonfantti;Mare RA Di;R Giovenardi
Check List 5 pages 819--829 (2009)

[5] Chromosomal evolution of South American frugivorous butterflies in the Satyroid clade (Nymphalidae: Charaxinae, Morphinae and Satyrinae).
by Jr KS Brown;AVL Freitas;BV Schoultz;AO Saura;A Saura
Biological Journal of the Linnean Society 92 pages 467--481 (2007)
10.1111/j.1095-8312.2007.00872.x

[6] 10.1111/j.1095-8312.2007.00872.x

[7] by B D’Abrera

[8] Borboletas (Papilionoidea e Hesperioidea) de Guarapuava e arredores, Paraná, Brasil: um inventário com base em 63 anos de registros.
by DR Dolibaina;OHH Mielke;MM Casagrande
Biota Neotropica 11 pages 343--354 (2011)
10.1590/s1676-06032011000100031

[9] 10.1590/s1676-06032011000100031

[10] On the frequency of butterflies in eastern Brazil, with a list of butterfly fauna of Poços de Caldas, Minas Gerais.
by H Ebert
Journal of the Lepidopterists' Society 23 pages 1--48 (1969)
144150

[11] 144150

[12] Beiträge zur Kenntnis der Insektenfauna Boliviens XIX. Lepidoptera III. Satyridae.
by W Forster
Veröffentlichungen der zoologischen Staatssammlung München 8 pages 51--188 (1964)
113851

[13] 113851

[14] A new species of Yphthimoides (Nymphalidae, Satyrinae) from Southeastern Brazil.
by AVL Freitas
Journal of the Lepidopterists' Society 58 pages 7--12 (2004)
113839

[15] 113839

[16] A new species of Moneuptychia Forster (Lepidoptera: Satyrinae, Euptychiina) from the highlands of southeastern Brazil.
by AVL Freitas
Neotropical Entomology 36 pages 919--925 (2007)
10.1590/s1519-566x2007000600014

[17] 10.1590/s1519-566x2007000600014

[18] Description of genus Guaianaza for “Euptychia” pronophila (Lepidoptera: Nymphalidae: Satyrinae) with a description of the immature stages.
by AVL Freitas;C Peña
Zootaxa 1163 pages 49--59 (2006)

[19] A new species of Moneuptychia Forster (Lepidoptera: Satyrinae: Euptychiina) from central Brazil.
by AVL Freitas;EO Emery;OHH Mielke
Neotropical Entomology 39 pages 83--90 (2010)
10.1590/s1519-566x2010000100011

[20] 10.1590/s1519-566x2010000100011

[21] A new genus and species of Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from southern Brazil.
by AVL Freitas;OHH Mielke;A Moser;KL Silva-Brandão;CA Iserhard
Neotropical Entomology 40 pages 231--237 (2011)
10.1590/S1519-566X2011000200012

[22] 10.1590/S1519-566X2011000200012

[23] Zoologia, Curitiba.
by AVL Freitas;EP Barbosa;JP Santos;OHH Mielke

[24] Diversidade de Lepidoptera (Papilionoidea e Hesperioidea) em dois fragmentos de floresta no município de Frederico Westphalen, Rio Grande do Sul, Brasil.
by R Giovenardi;RA Di Mare;J Sponchiado;S Roani;FAF Jacomassa;AB Jung;MA Porn
Revista Brasileira de Entomologia 52 pages 599--605 (2008)
10.1590/S0085-56262008000400010

[25] 10.1590/S0085-56262008000400010

[26] Lista de espécies de borboletas (Lepidoptera, Papilionoidea e Hesperioidea) da região do vale do rio Maquiné, Rio Grande do Sul, Brasil.
by CA Iserhard;HP Romanowski
Revista Brasileira de Zoologia 21 pages 649--662 (2004)
10.1590/s0101-81752004000300027

[27] 10.1590/s0101-81752004000300027

[28] Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) ocorrentes em diferentes ambientes na Floresta Ombrófila Mista e nos Campos de Cima da Serra do Rio Grande do Sul, Brasil.
by CA Iserhard;MT Quadros;HP Romanowski;MS Mendonça
Biota Neotropica 10 pages 309--320 (2010)
10.1590/s1676-06032010000100026

[29] 10.1590/s1676-06032010000100026

[30] by JA Kochalka;D Torres;B Garcete;C Aguilar

[31] Papilionoidea (Lepidoptera) de Pelotas e seus arredores, Rio Grande do Sul, Brasil.
by CP Krüger;EJE Silva
Entomologia y Vectores 10 pages 31--45 (2003)

[32] by G Lamas

[33] From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera: Nymphalidae): history, progress and prospects.
by MA Marín;C Peña;AVL Freitas;N Wahlberg;SI Uribe
Neotropical Entomology 40 pages 1--13 (2011)
10.1590/s1519-566x2011000100001

[34] 10.1590/s1519-566x2011000100001

[35] Systematics and evolutionary history of butterflies in the ‘‘Taygetis clade’'(Nymphalidae: Satyrinae: Euptychiina): Towards a better understanding of Neotropical biogeography.
by PF Matos-Maraví;C Peña;KR Willmott;AVL Freitas;N Wahlberg
Molecular Phylogenetics and Evolution 66 pages 54--68 (2013)
10.1016/j.ympev.2012.09.005

[36] 10.1016/j.ympev.2012.09.005

[37] by DL Murray

[38] Immature stages and biology of Taygetis Hübner (Lepidoptera: Nymphalidae).
by DL Murray
Proceedings of the Entomological Society of Washington 103 pages 932--945 (2001b)

[39] A catalogue of the type specimens of Nymphalidae deposited in the collection of the Staatliches Museum für Tierkunde Dresden (Insecta: Lepidoptera: Rhopalocera).
by YP Nekrutenko
Entomologische Abhandlungen Museum für Tierkunde Dresden 59 pages 325--403 (2001)

[40] Bulletin de la Sociéte Entomologique de Mulhouse
by EV Niculescu

[41] Sobre a morfologia do penis em Lepidoptera.
by J Oiticica-Filho
Boletim do Museu Nacional, Rio de Janeiro, n. s., Zoologia 50 pages 1--79 (1946)

[42] Nymphalidae, Papilionidae e Pieridae (Lepidoptera: Papilionoidea) da Serra do Sudoeste do Rio Grande do Sul, Brasil.
by ALG Paz;HP Romanowski;ABB Morais
Biota Neotropica 8 pages 141--149 (2008)
10.1590/s1676-06032008000100017

[43] 10.1590/s1676-06032008000100017

[44] Occurrence of fruit-feeding butterflies (Lepidoptera: Nymphalidae) in a fragment of Araucaria Moist Forest in Rio Grande do Sul State, Brazil.
by VS Pedrotti;MP Barros;HP Romanowski;CA Iserhard
Biota Neotropica 11 pages 385--390 (2011)
10.1590/S1676-06032011000100036

[45] 10.1590/S1676-06032011000100036

[46] Revision of the butterfly genus Forsterinaria Gray, 1973 (Lepidoptera: Nymphalidae, Satyrinae).
by C Peña;G Lamas
Revista Peruana de Biologia 12 pages 5--48 (2005)

[47] Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae).
by C Peña;S Nylin;AVL Freitas;N Wahlberg
Zoologica Scripta 39 pages 243--258 (2010)
10.1111/j.1463-6409.2010.00421.x

[48] 10.1111/j.1463-6409.2010.00421.x

[49] Ninfalídeos (Lepidoptera, Nymphalidae) ocorrentes no norte da Planície Costeira do Rio Grande do Sul, Brasil.
by FC Quadros;AL Dorneles;E Corseuil
Biociências 12 pages 147--164 (2004)

[50] The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape.
by DB Ribeiro;R Batista;PI Prado;Jr. KS Brown;AVL Freitas
Biodiversity and Conservation 21 pages 811--827 (2012)
10.1007/s10531-011-0222-x

[51] 10.1007/s10531-011-0222-x

[52] Fruit-feeding butterflies guide of subtropical Atlantic Forest and Araucaria Moist Forest in State of Rio Grande do Sul, Brazil.
by JP Santos;CA Iserhard;MO Teixeira;HP Romanowski
Biota Neotropica 11 pages 253--274 (2011)
10.1590/s1676-06032011000300022

[53] 10.1590/s1676-06032011000300022

[54] Borboletas frugívoras (Lepidoptera: Nymphalidae) de uma área urbana (Área de Proteção Especial Manancial Cercadinho) em Belo Horizonte, Minas Gerais, Brasil.
by ARM Silva;CO Castro;PO Mafia;MOC Mendonça;TCC Alves;MV Beirão
Biota Neotropica 12 pages 292--297 (2012)
10.1590/S1676-06032012000300028

[55] 10.1590/S1676-06032012000300028

[56] Preliminary analysis of the diurnal Lepidoptera fauna of the Três Picos State Park, Rio de Janeiro, Brazil, with a note on Parides ascanius (Cramer, 1775).
by A Soares;JMS Bizarro;CB Bastos;N Tangerini;NA Silva;AS Silva;GB Silva
Tropical Lepidoptera Research 21 pages 66--79 (2011)

[57] Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (São Paulo).
by M Uehara-Prado;AVL Freitas;RB Francini;Jr KS Brown
Biota Neotropica 4 pages 1--5 (2004)
10.1590/S1676-06032004000100007

[58] 10.1590/S1676-06032004000100007

[59] Illustrated Lists of American Butterflies.
by AD Warren;KJ Davis;EM Stangeland;JP Pelham;NV Grishin

[60] Stegosatyrus, a new genus of Euptychiina from the grasslands of neotropical realm (Lepidoptera: Nymphalidae: Satyrinae).
by T Zacca;OHH Mielke;TW Pyrcz;MM Casagrande;AVL Freitas;P Boyer
Zootaxa 3682 pages 331--350 (2013)
10.11646/zootaxa.3682.2.7

[61] 10.11646/zootaxa.3682.2.7

Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) de Matas Paludosas e Matas de Restinga da Planície Costeira da região Sul do Brasil.

Arquivos de Entomologia, série A

Butterflies (Lepidoptera: Papilionoidea e Hesperioidea) from two forest fragments in northern Rio Grande do Sul, Brazil.

Chromosomal evolution of South American frugivorous butterflies in the Satyroid clade (Nymphalidae: Charaxinae, Morphinae and Satyrinae).

Borboletas (Papilionoidea e Hesperioidea) de Guarapuava e arredores, Paraná, Brasil: um inventário com base em 63 anos de registros.

Zoologia, Curitiba.

Diversidade de Lepidoptera (Papilionoidea e Hesperioidea) em dois fragmentos de floresta no município de Frederico Westphalen, Rio Grande do Sul, Brasil.

Lista de espécies de borboletas (Lepidoptera, Papilionoidea e Hesperioidea) da região do vale do rio Maquiné, Rio Grande do Sul, Brasil.

Borboletas (Lepidoptera: Papilionoidea e Hesperioidea) ocorrentes em diferentes ambientes na Floresta Ombrófila Mista e nos Campos de Cima da Serra do Rio Grande do Sul, Brasil.

Papilionoidea (Lepidoptera) de Pelotas e seus arredores, Rio Grande do Sul, Brasil.

From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera: Nymphalidae): history, progress and prospects.

Systematics and evolutionary history of butterflies in the ‘‘Taygetis clade’'(Nymphalidae: Satyrinae: Euptychiina): Towards a better understanding of Neotropical biogeography.

Immature stages and biology of Taygetis Hübner (Lepidoptera: Nymphalidae).

A catalogue of the type specimens of Nymphalidae deposited in the collection of the Staatliches Museum für Tierkunde Dresden (Insecta: Lepidoptera: Rhopalocera).

Bulletin de la Sociéte Entomologique de Mulhouse

Nymphalidae, Papilionidae e Pieridae (Lepidoptera: Papilionoidea) da Serra do Sudoeste do Rio Grande do Sul, Brasil.

Occurrence of fruit-feeding butterflies (Lepidoptera: Nymphalidae) in a fragment of Araucaria Moist Forest in Rio Grande do Sul State, Brazil.

Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae).

Ninfalídeos (Lepidoptera, Nymphalidae) ocorrentes no norte da Planície Costeira do Rio Grande do Sul, Brasil.

The importance of small scales to the fruit-feeding butterfly assemblages in a fragmented landscape.

Fruit-feeding butterflies guide of subtropical Atlantic Forest and Araucaria Moist Forest in State of Rio Grande do Sul, Brazil.

Borboletas frugívoras (Lepidoptera: Nymphalidae) de uma área urbana (Área de Proteção Especial Manancial Cercadinho) em Belo Horizonte, Minas Gerais, Brasil.

Preliminary analysis of the diurnal Lepidoptera fauna of the Três Picos State Park, Rio de Janeiro, Brazil, with a note on Parides ascanius (Cramer, 1775).

Guia das borboletas frugívoras da Reserva Estadual do Morro Grande e região de Caucaia do Alto, Cotia (São Paulo).

Illustrated Lists of American Butterflies.

The Taygetis ypthima species group (Lepidoptera, Nymphalidae, Satyrinae): taxonomy, variation and description of a new species

Authors