Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution

doi:10.3897/zookeys.417.7733

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Title	Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution
Author(s)	Igor Sokolov James Reddell David Kavanaugh
Journal	ZooKeys
ISSN	1313-2989
Volume	417
Pages	71--101
DOI	10.3897/zookeys.417.7733
Year	2014
Publisher	Pensoft Publishers
Citation

...

Taxonomic review of the tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution.
by GE Ball;D Shpeley
Zootaxa 1099 (2005)
A taxonomic review of the genus Apenes LeConte (Coleoptera: Carabidae: Lebiini) in the West Indies, with descriptions of new species and notes about classification and biogeography.
by GE Ball;D Shpeley
Annals of the Carnegie Museum 78 pages 79--191 (2009)
- 10.2992/007.078.0201
A new genus and species of stygobiontic dryopid beetle, Stygoparnus comalensis (Coleoptera: Dryopidae), from Comal Springs, Texas.
by CB Barr;PJ Spangler
Proceedings of the Biological Society of Washington 105 pages 40--54 (1992)
- 85679
Revision of Rhadine LeConte (Coleoptera, Carabidae). I. The subterranea group.
by TC Barr
American Museum Novitates 2539 pages 1--30 (1974)
- 2246/5413
Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species.
by TC Barr
Special Bulletin Japanese Society of Coleopterology 4 pages 239--248 (1995)
Fourth supplement 1933 to 1938 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John. D.
by RE Blackwelder
Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico.
by Y Bousquet
Zookeys 245 pages 1--1722 (2012)
- 10.3897/zookeys.245.3416
New and little-known guests of the Texan legionary ants.
by CT Brues
The American Naturalist, an Illustrated Magazine of Natural History 36 pages 365--378 (1902)
Biogeographic affinities of pselaphid beetles of the Eastern United States.
by CE Carlton
Florida Entomologist 73 pages 570--579 (1990)
A review of the North American Bembidiinae. In: Casey TL (Ed) Memoirs on the Coleoptera. VIII.
by TL Casey
The Pselaphidae of Texas caves (Coleoptera)
by DS Chandler
Texas Memorial Museum, Speleological Monographs 3 pages 241--253 (1992)
A review of the ant-like litter beetles found in Texas caves (Coleoptera: Staphylinidae: Pselaphinae).
by DS Chandler;JR Reddell
Texas Memorial Museum, Speleological Monographs 5 pages 115--128 (2001)
New cave Pselaphinae and records from Texas, with a discussion of the relationships and distributions of the Texas troglobitic Pselaphinae (Coleoptera: Staphylinidae: Pselaphinae).
by DS Chandler;JR Reddell;P Paquin
Texas Memorial Museum Speleological Monographs 7 pages 125--140 (2009)
The North American obligate cave fauna: regional patterns.
by DC Culver;M Christman;WE Elliott;III H Hobbs;JR Reddell
Biodiversity and Conservation 12 pages 441--468 (2003)
The mid-latitude biodiversity ridge in terrestrial cave fauna.
by DC Culver;L Deharveng;A Bedos;JJ Lewis;M Madden;JR Reddell;B Sket;P Trontelj;D White
Ecography 29 pages 120--128 (2006)
A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera).
by TL Erwin;GN House
The Coleopoterists Bulletin 32 pages 231--256 (1978)
Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini).
by PM Giachino;D Vailati
Biodiversity Journal 1 pages 1--112 (2011)
- 9788890492983
First records of 13 mammalian species within the Southwestern Cross Timbers Region of Texas.
by JR Goetze;AD Nelson
Occasional Papers of Texas Tech University 284 pages 1--6 (2009)
Texas Agriculture Experiment Station Miscellaneous Publication
by FW Gould
Late Cretaceous and Cenozoic History of North American vegetation (north of Mexico).
by A Graham
Checklist of the vascular plants of Texas.
by SL Hatch;KN Gandhi;LE Brown
Texas Agriculture Experiment Station Miscellaneous Publication 1655 pages 1--158 (1990)
Systematics, speciation, and distribution of the subterranean amphipod genus Stygonectes (Gammaridae).
by JR Holsinger
Bulletin of the United States National Museum 259 pages 1--176 (1967)
- 10.5479/si.03629236.259.1
- 106013
Distributional patterns of Texas fresh-water fishes.
by C Hubbs
The Southwestern Naturalist 2 pages 89--104 (1957)
Description of a new genus and species of stygobiontic diving beetle, Psychopomporus felipi Jean, Telles, and Miller (Coleoptera: Dytiscidae: Hydroporinae), from the Edwards-Trinity aquifer system of Texas, USA.
by A Jean;ND Telles;JR Gibson;D Foley;KB Miller
The Coleopterists Bulletin 66 pages 105--110 (2012)
- 10.1649/072.066.0202
Les Bembidiides endogés (Col. Carabidae). Monographie d’une lignée gondwanienne.
by R Jeannel
Revue Française d’Entomologie 3 pages 241--339 (1937)
Monographie des "Anillini", bembidiides endogés [Coleoptera Trechidae].
by R Jeannel
Mémoires du Muséum National d'Histoire Naturelle (Série A) Zoologie 28 pages 33--204 (1963a)
Supplément à la monographie des Anillini. Sur quelques espèces nouvelles de l'Amérique du Nord.
by R Jeannel
Revue Française d'Entomologie 30 pages 145--152 (1963b)
Notes on the classification of the Carabidae of the United States.
by JL LeConte
Transactions of the American Philosophical Society 10 pages 363--403 (1853)
Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves.
by JP Ledford;P Paquin;JC Cokendolpher;J Campbell;C Griswold
Zookeys 167 pages 1--102 (2012)
- 10.3897/zookeys.167.1833
- 3272638
- 22363201
Catalogue of the Coleoptera of America, north of Mexico
by CW Leng
North American stygobiontic diving beetles (Coleoptera: Dytiscidae: Hydroporinae) with description of Ereboporus naturaconservatus Miller, Gibson and Alarie, new genus and species, from Texas, U.S.A.
by KB Miller;JR Gibson;Y Alarie
The Coleopterists Bulletin 63 pages 191--202 (2009)
- 10.1649/1124.1
The first troglobitic weevil (Coleoptera: Curculionidae) in North America? Description of a new eyeless species of Lymantes Schoenherr from Central Texas caves.
by P Paquin;RS Anderson
Texas Memorial Museum Speleological Monographs 7 pages 115--123 (2009)
The power and perils of ‘molecular taxonomy’: a case study of eyeless and endangered Cicurina (Araneae: Dictynidae) from Texas caves.
by P Paquin;M Hedin
Molecular Ecology 13 pages 3239--3255 (2004)
The Caves and Karst of Texas
by JR Reddell
A new species of troglobitic Rhadine (Coleoptera: Carabidae) from Texas.
by JR Reddell;JC Cokendolpher
Texas Memorial Museum Speleological Monographs 5 pages 109--114 (2001)
New species and records of cavernicole Rhadine (Coleoptera: Carabidae) from Camp Bullis, Texas.
by JR Reddell;JC Cokendolpher
Texas Memorial Museum Speleological Monographs 6 pages 153--162 (2004)
A new species of troglobitic Rhadine (Coleoptera: Carabidae) from Hays County, Texas.
by JR Reddell;N Dupérré
Texas Memorial Museum Speleological Monographs 7 pages 111--114 (2009)
Hydrologic Investigations Atlas 730-E
by PD Ryder
Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U.S.A. (Coleoptera: Carabidae: Trechinae: Bembidiini).
by IM Sokolov
Insecta Mundi 164 pages 1--14 (2011)
Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini).
by IM Sokolov
Annals of Carnegie Museum 81 pages 61--71 (2012)
- 10.2992/007.081.0104
A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistics analysis and some notes on the evolution of the genus.
by IM Sokolov
ZooKeys 352 pages 51--92 (2013)
- 10.3897/zookeys.352.6052
Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U.S.A.
by IM Sokolov;CE Carlton
Zootaxa 1740 pages 37--44 (2008)
Species of Anillinus (Coleoptera: Carabidae: Trechinae) described from Brazil and their relation to North American representatives of the genus.
by IM Sokolov;CE Carlton
Coleopterists Bulletin 66 pages 245--249 (2012)
Review of Anillinus, with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Carabidae: Trechinae: Bembidiini).
by IM Sokolov;CE Carlton;JF Cornell
The Coleopterists Bulletin 58 pages 185--233 (2004)
- 10.1649/611
A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region.
by IM Sokolov;LE Watrous
The Coleopterists Bulletin 62 pages 537--543 (2008)
- 10.1649/1114.1
A new genus and species of stygobiontic dytiscid beetle, Comaldessus stygius (Coleoptera: Dytiscidae: Bidessini), from Comal Springs, Texas.
by PJ Spangler;CB Barr
Insecta Mundi 9 pages 301--308 (1995)
50 CFR part 17, RIN 1018-AF33; Endangered and threatened wildlife and plants; Final rule to list nine Bexar County, Texas invertebrates species as endangered [prepared by C. Longacre].
by
Federal Register 65 pages 81419--81433 (2000)
A new subterranean aquatic beetle from Texas (Coleoptera: Dytiscidae: Hydroporinae).
by FN Young;G Longley
Annals of the Entomological Society of America 69 pages 787--792 (1976)

Only known citations between publications in BioNames are listed here.

[1] Taxonomic review of the tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution.
by GE Ball;D Shpeley
Zootaxa 1099 (2005)

[2] A taxonomic review of the genus Apenes LeConte (Coleoptera: Carabidae: Lebiini) in the West Indies, with descriptions of new species and notes about classification and biogeography.
by GE Ball;D Shpeley
Annals of the Carnegie Museum 78 pages 79--191 (2009)
10.2992/007.078.0201

[3] 10.2992/007.078.0201

[4] A new genus and species of stygobiontic dryopid beetle, Stygoparnus comalensis (Coleoptera: Dryopidae), from Comal Springs, Texas.
by CB Barr;PJ Spangler
Proceedings of the Biological Society of Washington 105 pages 40--54 (1992)
85679

[5] 85679

[6] Revision of Rhadine LeConte (Coleoptera, Carabidae). I. The subterranea group.
by TC Barr
American Museum Novitates 2539 pages 1--30 (1974)
2246/5413

[7] 2246/5413

[8] Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from southeastern United States, with description of a new genus and two new species.
by TC Barr
Special Bulletin Japanese Society of Coleopterology 4 pages 239--248 (1995)

[9] Fourth supplement 1933 to 1938 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John. D.
by RE Blackwelder

[10] Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico.
by Y Bousquet
Zookeys 245 pages 1--1722 (2012)
10.3897/zookeys.245.3416

[11] 10.3897/zookeys.245.3416

[12] New and little-known guests of the Texan legionary ants.
by CT Brues
The American Naturalist, an Illustrated Magazine of Natural History 36 pages 365--378 (1902)

[13] Biogeographic affinities of pselaphid beetles of the Eastern United States.
by CE Carlton
Florida Entomologist 73 pages 570--579 (1990)

[14] A review of the North American Bembidiinae. In: Casey TL (Ed) Memoirs on the Coleoptera. VIII.
by TL Casey

[15] The Pselaphidae of Texas caves (Coleoptera)
by DS Chandler
Texas Memorial Museum, Speleological Monographs 3 pages 241--253 (1992)

[16] A review of the ant-like litter beetles found in Texas caves (Coleoptera: Staphylinidae: Pselaphinae).
by DS Chandler;JR Reddell
Texas Memorial Museum, Speleological Monographs 5 pages 115--128 (2001)

[17] New cave Pselaphinae and records from Texas, with a discussion of the relationships and distributions of the Texas troglobitic Pselaphinae (Coleoptera: Staphylinidae: Pselaphinae).
by DS Chandler;JR Reddell;P Paquin
Texas Memorial Museum Speleological Monographs 7 pages 125--140 (2009)

[18] The North American obligate cave fauna: regional patterns.
by DC Culver;M Christman;WE Elliott;III H Hobbs;JR Reddell
Biodiversity and Conservation 12 pages 441--468 (2003)

[19] The mid-latitude biodiversity ridge in terrestrial cave fauna.
by DC Culver;L Deharveng;A Bedos;JJ Lewis;M Madden;JR Reddell;B Sket;P Trontelj;D White
Ecography 29 pages 120--128 (2006)

[20] A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera).
by TL Erwin;GN House
The Coleopoterists Bulletin 32 pages 231--256 (1978)

[21] Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini).
by PM Giachino;D Vailati
Biodiversity Journal 1 pages 1--112 (2011)
9788890492983

[22] 9788890492983

[23] First records of 13 mammalian species within the Southwestern Cross Timbers Region of Texas.
by JR Goetze;AD Nelson
Occasional Papers of Texas Tech University 284 pages 1--6 (2009)

[24] Texas Agriculture Experiment Station Miscellaneous Publication
by FW Gould

[25] Late Cretaceous and Cenozoic History of North American vegetation (north of Mexico).
by A Graham

[26] Checklist of the vascular plants of Texas.
by SL Hatch;KN Gandhi;LE Brown
Texas Agriculture Experiment Station Miscellaneous Publication 1655 pages 1--158 (1990)

[27] Systematics, speciation, and distribution of the subterranean amphipod genus Stygonectes (Gammaridae).
by JR Holsinger
Bulletin of the United States National Museum 259 pages 1--176 (1967)
10.5479/si.03629236.259.1
106013

[28] 10.5479/si.03629236.259.1

[29] 106013

[30] Distributional patterns of Texas fresh-water fishes.
by C Hubbs
The Southwestern Naturalist 2 pages 89--104 (1957)

[31] Description of a new genus and species of stygobiontic diving beetle, Psychopomporus felipi Jean, Telles, and Miller (Coleoptera: Dytiscidae: Hydroporinae), from the Edwards-Trinity aquifer system of Texas, USA.
by A Jean;ND Telles;JR Gibson;D Foley;KB Miller
The Coleopterists Bulletin 66 pages 105--110 (2012)
10.1649/072.066.0202

[32] 10.1649/072.066.0202

[33] Les Bembidiides endogés (Col. Carabidae). Monographie d’une lignée gondwanienne.
by R Jeannel
Revue Française d’Entomologie 3 pages 241--339 (1937)

[34] Monographie des "Anillini", bembidiides endogés [Coleoptera Trechidae].
by R Jeannel
Mémoires du Muséum National d'Histoire Naturelle (Série A) Zoologie 28 pages 33--204 (1963a)

[35] Supplément à la monographie des Anillini. Sur quelques espèces nouvelles de l'Amérique du Nord.
by R Jeannel
Revue Française d'Entomologie 30 pages 145--152 (1963b)

[36] Notes on the classification of the Carabidae of the United States.
by JL LeConte
Transactions of the American Philosophical Society 10 pages 363--403 (1853)

[37] Systematics, conservation and morphology of the spider genus Tayshaneta (Araneae, Leptonetidae) in Central Texas Caves.
by JP Ledford;P Paquin;JC Cokendolpher;J Campbell;C Griswold
Zookeys 167 pages 1--102 (2012)
10.3897/zookeys.167.1833
3272638
22363201

[38] 10.3897/zookeys.167.1833

[39] 3272638

[40] 22363201

[41] Catalogue of the Coleoptera of America, north of Mexico
by CW Leng

[42] North American stygobiontic diving beetles (Coleoptera: Dytiscidae: Hydroporinae) with description of Ereboporus naturaconservatus Miller, Gibson and Alarie, new genus and species, from Texas, U.S.A.
by KB Miller;JR Gibson;Y Alarie
The Coleopterists Bulletin 63 pages 191--202 (2009)
10.1649/1124.1

[43] 10.1649/1124.1

[44] The first troglobitic weevil (Coleoptera: Curculionidae) in North America? Description of a new eyeless species of Lymantes Schoenherr from Central Texas caves.
by P Paquin;RS Anderson
Texas Memorial Museum Speleological Monographs 7 pages 115--123 (2009)

[45] The power and perils of ‘molecular taxonomy’: a case study of eyeless and endangered Cicurina (Araneae: Dictynidae) from Texas caves.
by P Paquin;M Hedin
Molecular Ecology 13 pages 3239--3255 (2004)

[46] The Caves and Karst of Texas
by JR Reddell

[47] A new species of troglobitic Rhadine (Coleoptera: Carabidae) from Texas.
by JR Reddell;JC Cokendolpher
Texas Memorial Museum Speleological Monographs 5 pages 109--114 (2001)

[48] New species and records of cavernicole Rhadine (Coleoptera: Carabidae) from Camp Bullis, Texas.
by JR Reddell;JC Cokendolpher
Texas Memorial Museum Speleological Monographs 6 pages 153--162 (2004)

[49] A new species of troglobitic Rhadine (Coleoptera: Carabidae) from Hays County, Texas.
by JR Reddell;N Dupérré
Texas Memorial Museum Speleological Monographs 7 pages 111--114 (2009)

[50] Hydrologic Investigations Atlas 730-E
by PD Ryder

[51] Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U.S.A. (Coleoptera: Carabidae: Trechinae: Bembidiini).
by IM Sokolov
Insecta Mundi 164 pages 1--14 (2011)

[52] Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini).
by IM Sokolov
Annals of Carnegie Museum 81 pages 61--71 (2012)
10.2992/007.081.0104

[53] 10.2992/007.081.0104

[54] A new genus and eight new species of the subtribe Anillina (Carabidae, Trechinae, Bembidiini) from Mexico, with a cladistics analysis and some notes on the evolution of the genus.
by IM Sokolov
ZooKeys 352 pages 51--92 (2013)
10.3897/zookeys.352.6052

[55] 10.3897/zookeys.352.6052

[56] Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U.S.A.
by IM Sokolov;CE Carlton
Zootaxa 1740 pages 37--44 (2008)

[57] Species of Anillinus (Coleoptera: Carabidae: Trechinae) described from Brazil and their relation to North American representatives of the genus.
by IM Sokolov;CE Carlton
Coleopterists Bulletin 66 pages 245--249 (2012)

[58] Review of Anillinus, with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Carabidae: Trechinae: Bembidiini).
by IM Sokolov;CE Carlton;JF Cornell
The Coleopterists Bulletin 58 pages 185--233 (2004)
10.1649/611

[59] 10.1649/611

[60] A new species and the first record of the genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark region.
by IM Sokolov;LE Watrous
The Coleopterists Bulletin 62 pages 537--543 (2008)
10.1649/1114.1

[61] 10.1649/1114.1

[62] A new genus and species of stygobiontic dytiscid beetle, Comaldessus stygius (Coleoptera: Dytiscidae: Bidessini), from Comal Springs, Texas.
by PJ Spangler;CB Barr
Insecta Mundi 9 pages 301--308 (1995)

[63] 50 CFR part 17, RIN 1018-AF33; Endangered and threatened wildlife and plants; Final rule to list nine Bexar County, Texas invertebrates species as endangered [prepared by C. Longacre].
by
Federal Register 65 pages 81419--81433 (2000)

[64] A new subterranean aquatic beetle from Texas (Coleoptera: Dytiscidae: Hydroporinae).
by FN Young;G Longley
Annals of the Entomological Society of America 69 pages 787--792 (1976)

Fourth supplement 1933 to 1938 (inclusive) to the Leng catalogue of Coleoptera of America, north of Mexico. John. D.

Biogeographic affinities of pselaphid beetles of the Eastern United States.

A review of the ant-like litter beetles found in Texas caves (Coleoptera: Staphylinidae: Pselaphinae).

The North American obligate cave fauna: regional patterns.

The mid-latitude biodiversity ridge in terrestrial cave fauna.

A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera).

First records of 13 mammalian species within the Southwestern Cross Timbers Region of Texas.

Texas Agriculture Experiment Station Miscellaneous Publication

Late Cretaceous and Cenozoic History of North American vegetation (north of Mexico).

Checklist of the vascular plants of Texas.

Distributional patterns of Texas fresh-water fishes.

Monographie des "Anillini", bembidiides endogés [Coleoptera Trechidae].

Supplément à la monographie des Anillini. Sur quelques espèces nouvelles de l'Amérique du Nord.

Catalogue of the Coleoptera of America, north of Mexico

The power and perils of ‘molecular taxonomy’: a case study of eyeless and endangered Cicurina (Araneae: Dictynidae) from Texas caves.

The Caves and Karst of Texas

Hydrologic Investigations Atlas 730-E

Species of Anillinus (Coleoptera: Carabidae: Trechinae) described from Brazil and their relation to North American representatives of the genus.

50 CFR part 17, RIN 1018-AF33; Endangered and threatened wildlife and plants; Final rule to list nine Bexar County, Texas invertebrates species as endangered [prepared by C. Longacre].

Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution

Authors