Yoshimatsu, Toriumi & Dodge 2004. Morphology and taxonomy of five marine sand-dwelling Thecadinium species (Dinophyceae) from Japan, including four new species: Thecadinium arenarium sp. nov., Thecadinium ovatum sp. nov., Thecadinium striatum sp. nov. and Thecadinium yashimaense. sp. nov. Phycological Research, 52(3), September 2004: 211-223