Ratcliffe 1980. New species of Coprini (Coleoptera: Scarabaeidae: Scarabaeinae) taken from the pelage of three toed sloths (Bradypus tridactylus L.) (Edentata: Bradypodidae) in central Amazonia with a brief commentary on scarab-sloth relationships. Coleopterists Bulletin, 34(4) 1980: 337-350